Madonna has performed "Human Nature" three times on tour. In her 2001 Drowned World Tour she performed the song while slow-riding on a mechanical bull. During the performance she was dressed in blue jeans and a shirt with a U.S. flag design on it. The song was accompanied by a vocoder, which played the lines "Did I say something wrong?" and "Oops, I didn't know I couldn't talk about sex." Madonna performed the song again at her 2008 Sticky & Sweet Tour, with an electric guitar, also adding vocoder voice effects on the background harmonies, with a video backdrop of American singer Britney Spears trapped in an elevator and tried to get out. At the end of the performance, the doors opened to reveal Spears saying the phrase "It's Britney, bitch", from her song "Gimme More" (2007). On November 6, 2008 in the Los Angeles show at Dodger Stadium, Spears joined Madonna onstage halfway through the performance.
In 2012, Madonna performed the song on her MDNA Tour as part of the Masculine/Feminine segment of the show. She performed the song while her dancers moved mirrors around her, as she removed articles of clothing. At her concert in Istanbul on June 7, 2012, Madonna briefly exposed her right breast to the audience, before covering it up again. This prompted some criticism about her age, with one critic, Sophie Wilkinson, asking, "at 54, is she too old to be getting a boob out?" The Sun notes, "Her actions may have been offensive to some members of her audience, as Turkey is a traditionally Islamic country, with a largely Muslim population" before noting that "Perhaps fans shouldn't have been surprised - as she did warn them, writing No Fear on her back." On the other hand, Amy Odell from BuzzFeed said that criticizing the gesture based on her age send "a very archaic message", adding that "Women who are 53 still have breasts and... sex drives!".
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