In Video Games
After the ruler of the other dimensional realm Outworld, Shao Kahn, conquered the realm of Edenia and merged it with his own, he decided he would keep the former king's daughter Kitana alive and raise her as his own. Though she grew up knowing nothing of her origin, the emperor nonetheless feared that one day Kitana would discover her true parentage and turn against him, and so he ordered Shang Tsung to bring to life a more vicious and loyal version of Kitana that could take her place if necessary. This was done by fusing her essence with a Tarkatan warrior from Baraka's race. The process was not a complete success, however, as the hybrid clone was disfigured with the hideous mouth of the Tarkata. Instead of replacing Kitana, as originally planned, Mileena would be used to spy on her and to ensure her allegiance to him, and so Shao Kahn introduced Mileena to Kitana as her supposedly lost twin sister. The two thus grew up together as his daughters and personal assassins, though Mileena (who has become a companion of Baraka) privately grew to harbor a great bitterness and jealousy towards Kitana, whom Kahn favored over her.
As Kahn feared, Kitana did finally find the truth about her past and so Mileena was ordered to keep close watch over her rival when Kitana secretly allied with Earthrealm warriors during the events of Mortal Kombat II (1993). Mileena, determined to stop her twin at any cost, is then murdered by Kitana, her soul descending into the Netherrealm. In Mortal Kombat: Shaolin Monks (2005), the beat'em up retelling of Mortal Kombat II, Mileena (voiced by Lita Lopez) fights against Liu Kang and Kung Lao alongside Jade and Kitana, but is defeated by the two Shaolin warriors and flees to the Wasteland to seek help from Goro. Damned to the Netherrealm after her death, Mileena swears fealty to its ruler Shinnok. During the events of Ultimate Mortal Kombat 3 (1996), Shao Kahn decides to resurrect Mileena so her skills as a vicious fighter would help him defeat Earth's chosen warriors, also magically granting her the ability to read Kitana's thoughts. Shinnok sees this as an opportunity to covertly monitor the events unfolding in Earthrealm and so he allows her to return to life. After Kahn is defeated, Mileena is summoned back to the Netherrealm. In Mortal Kombat Gold (1999), she assists in Shinnok's invasion of Edenia, but allows her sister to escape from a dungeon. Following Shinnok's defeat, Mileena comes to Kitana's palace and demands the power over Edenia to be shared with her but Kitana refuses and, when attacked, traps Mileena, locking her away in a dungeon.
Mileena remains imprisoned for years, until Onaga the Dragon King began his own invasion of Edenia, during which she is freed from her imprisonment by her ally Baraka. During the events of Mortal Kombat: Deception (2004), Mileena (voiced by Johanna AƱonuevo) is ordered by Onaga to pose as Kitana in order to confuse and misdirect his enemies. However, as Mileena begins the game's titular deception, she develops a hidden agenda and decides to take control of both the forces of Edenia and Onaga's own undead army for herself. In the game's story mode (Konquest), Mileena also trains the young Shujinko in the Outworld and fights against Jade. In Mortal Kombat: Armageddon (2006), Mileena seizes Shao Kahn's fortress, still under the guise of Kitana, and decides to continue her charade until the Edenian forces were corrupted enough to follow her under the true identity. Though at first confident that Outworld is hers to rule, Mileena is forced to reveal herself and surrender to the returning Shao Kahn when he mounts an offensive against the fortress to re-instate his rule over Outworld. The emperor then commands her to capture Shujinko, who would be used as a bargaining chip in gaining Onaga as an ally. Mileena, pretending to be Kitana, succeeds in capturing Shujinko, taking him to Shao Kahn's palace. But having tasted power for herself, she is no longer content with being his minion and still plans to get the throne of Edenia back for herself. She is later killed by Shang Tsung during the final all-out battle at the Soulnado.
Mileena returns in Mortal Kombat (2011), an alternative-timeline retelling of the original Mortal Kombat trilogy that brought some major changes to her character. In this game, Mileena (voiced by Karen Strassman, who was also hired for the next sequel) was created by Shang Tsung in the "Flesh Pit" and introduced during the second tournament, instead of many years earlier. Meant to be a loyal replacement for Kitana, she is physically and mentally damaged, her animalistic rage controlled only by Shao Kahn as she uses her lascivious behavior to lure victims and then slaughter and devour them. In the game's story mode, she serves as an opponent for Kitana, Jade, Kurtis Stryker and Kabal, and is referred to by Shao Kahn as his "true daughter"; when Kahn appears to be killed, his minions even discuss making her a new ruler of Outworld. Her new personality is much more immature (even infantile) than in the previous games, and even Kitana (in her uncanonical ending) is shown to be an unusually sympathic towards her.
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