Ground Beetles

Some articles on ground beetles, beetles, beetle:

Ground Beetle - Evolution and Systematics
... Ground beetles evolved in the latter Triassic, having separated from their closest relatives by 200 million years ago ... The closest living relatives of the ground beetles are the false ground beetles (Trachypachidae) and the wrinkled bark beetles (Rhysodidae) ... as subfamilies or as tribes incertae sedis, but more preferably they are united with the ground beetles in the superfamily Caraboidea ...
Trachypachidae
... The Trachypachidae (or sometimes false ground beetles) are a family of beetles that generally resemble small ground beetles, but that are distinguished by the large coxae of their rearmost legs ... Their habits are similar to those of the ground beetles they are usually found in the leaf litter of conifer forests ... as a family been placed in several different positions relative to other beetle families ...

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